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Blocking FOXO4, which inhibits the activation of P53, causes senescent cells to undergo apoptosis by P53 activation, without affecting healthy cells around them, and has anti-aging effects which may have potential applications in the treatment of numerous age-related diseases, as well as an adjunct to treatment for certain forms of cancer

The reason he calls this a hypothesis, is that even though many studies (some you will see below) have demonstrated this effect, there have not been enough large-scale randomized trials to definitively prove this out

ROS, produced by UV radiation, primarily target mitochondrial DNA (mtDNA), leading to mutations due to the lack of protective histones and mechanisms

J Pharm Pharmacol 113 Srinivasan K (2014) Antioxidant potential of spices and their active constituents