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With the rise of Arabidopsis as a study system, and notably genome sequencing, the pathway and its genes were characterized at the molecular genetic level, revealing (or confirming) that all four enzymes were encoded by more than one gene, thus explaining the presence of the enzyme activities in several compartments (Table 1)

In the study by Vilela et al

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This oxidative stress resulted in mitochondrial dysfunction, which was depicted by a loss of viability, or mitochondrial membrane potential (m), and loss of mitochondrial health, or mitochondrial mass